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SpoT–RelA regulatory logic and physiology under perturbed (p)ppGpp

Clarify the regulatory logic of SpoT and its interplay with RelA in (p)ppGpp control, and establish a comprehensive understanding of cellular physiology under perturbed (p)ppGpp conditions, including the validity of ppGpp homeostasis assumptions.

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Background

ppGpp is central to the bacterial stringent response; recent work relates ppGpp to elongation rate, yet multiple regulatory details remain unresolved.

Addressing this question would strengthen mechanistic links between sensing, ribosome activity, and proteome allocation under nutrient and translational perturbations.

References

On the modeling front, the recent advances connecting ppGpp dynamics to translation elongation in E.~coli demonstrate the power of hybrid approaches to bridge the power of coarse-grained models with the specific predictions of mechanistic descriptions. However, several open questions remain, such as the regulatory logic of SpoT and its interplay with RelA, and a full understanding of the physiology under perturbed ppGpp conditions, challenging long-standing assumptions about ppGpp homeostasis.

The Hands-On Growth Laws Theory Cookbook (2507.19194 - Droghetti et al., 25 Jul 2025) in Section 6, OUTLOOK