Sparse Crosscoders for diffing MoEs and Dense models

Abstract: Mixture of Experts (MoE) achieve parameter-efficient scaling through sparse expert routing, yet their internal representations remain poorly understood compared to dense models. We present a systematic comparison of MoE and dense model internals using crosscoders, a variant of sparse autoencoders, that jointly models multiple activation spaces. We train 5-layer dense and MoEs (equal active parameters) on 1B tokens across code, scientific text, and english stories. Using BatchTopK crosscoders with explicitly designated shared features, we achieve $\sim 87\%$ fractional variance explained and uncover concrete differences in feature organization. The MoE learns significantly fewer unique features compared to the dense model. MoE-specific features also exhibit higher activation density than shared features, whereas dense-specific features show lower density. Our analysis reveals that MoEs develop more specialized, focused representations while dense models distribute information across broader, more general-purpose features.