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Branching Diffusion with Interactions

Updated 11 July 2026
  • Branching Diffusion with Interactions is a stochastic framework where individuals diffuse and branch based on interaction-dependent rules, capturing complex population dynamics.
  • It incorporates various mechanisms such as rank-based interactions, mean-field effects, and catalytic branching to model both local and global behaviors.
  • Analytical methods like hydrodynamic limits, traveling wave analysis, and spine techniques provide insights into long-term system behavior and control.

Searching arXiv for recent and foundational papers on interacting branching diffusions to ground the article. Branching diffusion with interactions denotes a class of stochastic population models in which individuals move in continuous space, typically by diffusion, while branching, death, or selection events depend on the current population configuration rather than on independent per-particle mechanisms alone. Recent formulations represent the population either by a finite atomic measure on an Ulam–Harris tree or by a weighted empirical measure, and place the interaction in rank, local occupancy, empirical-law dependence, pairwise catalytic terms, or nonlinear competition (Claisse et al., 2024, Ocello, 16 Jan 2026, Demircigil et al., 13 May 2025). In this sense, the subject includes rank-based branching Brownian systems, McKean–Vlasov branching diffusions, mutually catalytic branching, spatially self-regulating diffusions, and multitype interacting branching processes, together with their hydrodynamic, mean-field, and genealogical limits.

1. State spaces, labeling schemes, and martingale formulations

A common representation uses Ulam–Harris labels to encode genealogies. In the controlled interacting branching-diffusion framework, the label set is

Ic={}    n1Nn,\mathcal I_c=\{\varnothing\}\;\cup\;\bigcup_{n\ge1}\mathbb N^n,

and the state space is

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},

endowed with the weak-* topology. A controlled branching-diffusion is then a càdlàg, adapted EE-valued process satisfying a martingale problem in which each labeled particle evolves by a diffusion, branches according to a configuration-dependent death rate, and produces kk offspring with probability pk(i,x,λ,a)p_k(i,x,\lambda,a) (Ocello, 16 Jan 2026).

An equivalent population-level notation appears in McKean–Vlasov branching diffusion. There one writes

Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},

with Kt\mathcal K_t the set of alive particles and μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t) the law of the whole branching process up to time tt. Each particle follows

dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,

dies at rate E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},0, and at death produces a random number of offspring with law E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},1 (Claisse et al., 2024).

For large but finite populations, weighted empirical measures are standard. In the rank-based Brownian system one records

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},2

where E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},3 is the set of alive labels and E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},4 is the rescaled population size (Demircigil et al., 13 May 2025). In mean-field interacting branching diffusions with logistic binary branching one likewise writes

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},5

so that total mass remains of order E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},6 under the E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},7-weighting (Fontbona et al., 2021).

Across these formulations, martingale problems provide the unifying analytic language. In the rank-based model, for each E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},8,

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},9

with a local generator EE0 and a martingale EE1 whose quadratic variation is EE2 (Demircigil et al., 13 May 2025). In the McKean–Vlasov setting, weak solutions are characterized by the martingale property of

EE3

for smooth test functions EE4, where EE5 is a nonlinear generator acting on the current configuration (Claisse et al., 2024).

2. Principal interaction mechanisms

One major class is rank-based interaction. In the “Go or Grow” system, the rank of a particle in a configuration EE6 is

EE7

The EE8 rightmost particles, with ranks EE9, are in the Grow regime and branch at rate kk0 each, while all others are in the Go regime, drift rightward with speed kk1, and do not branch (Demircigil et al., 13 May 2025). The motion between branching events is Brownian with piecewise-constant drift,

kk2

where kk3 indicates whether the particle has rank kk4 (Demircigil et al., 13 May 2025). The model is explicitly tied to the Go or Grow hypothesis for cells in a capillary tube moving upwards a chemical gradient.

A second class is mean-field interaction through smoothed empirical fields. In the quantitative mean-field model, each particle solves

kk5

with bounded Lipschitz kernels kk6, birth rate kk7, and death rate kk8 (Fontbona et al., 2021). The interaction therefore enters both the diffusion coefficients and the logistic mass regulation.

A third class is nonlinear dependence on the law of the whole branching process. In McKean–Vlasov branching diffusion, the coefficients kk9, the death rate pk(i,x,λ,a)p_k(i,x,\lambda,a)0, and the offspring laws pk(i,x,λ,a)p_k(i,x,\lambda,a)1 depend on pk(i,x,λ,a)p_k(i,x,\lambda,a)2, so the interaction is measure-valued and path-dependent (Claisse et al., 2024). Under boundedness, measurability, and Lipschitz conditions in path and measure under pk(i,x,λ,a)p_k(i,x,\lambda,a)3, strong well-posedness is obtained (Claisse et al., 2024).

Catalytic interaction produces a different mechanism. In finite-rate two-type mutually catalytic branching on a countable site space pk(i,x,λ,a)p_k(i,x,\lambda,a)4,

pk(i,x,λ,a)p_k(i,x,\lambda,a)5

where pk(i,x,λ,a)p_k(i,x,\lambda,a)6 and pk(i,x,λ,a)p_k(i,x,\lambda,a)7 implements negative correlation (Doering et al., 2011). In the discrete particle counterpart on pk(i,x,λ,a)p_k(i,x,\lambda,a)8, a type-1 particle at pk(i,x,λ,a)p_k(i,x,\lambda,a)9 branches at instantaneous rate Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},0, and symmetrically a type-2 particle branches at rate Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},1, where the catalyst is the local population of the opposite type (Fugenfirov et al., 2023).

Competition and self-regulation give another canonical structure. In the small-mass diffusion limit of a multitype locally self-regulating model,

Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},2

so migration, local carrying capacities, and cross-type competition are all explicit in the drift (Greven et al., 2015). A related one-dimensional competition limit is

Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},3

where Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},4 encodes net births minus deaths including competition (Ba et al., 2013).

Finally, interaction may act through the center of mass. In the dyadic branching diffusion with Ornstein–Uhlenbeck drift and mean-field attraction or repulsion,

Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},5

so the overall drift is the combination of Ornstein–Uhlenbeck confinement or expansion and pairwise mean-field attraction or repulsion (Englander et al., 2016).

3. Large-population, hydrodynamic, and scaling limits

The rank-based Go or Grow system admits a hydrodynamic limit as Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},6. Writing Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},7, defining the cumulative tail

Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},8

and weighting individuals by Zt=kKtδ(k,Xtk),Z_t=\sum_{k\in\mathcal K_t}\delta_{(k,X^k_t)},9, the limit is the nonlinear integro-PDE

Kt\mathcal K_t0

or, in terms of Kt\mathcal K_t1,

Kt\mathcal K_t2

with tightness and convergence established under finite third-moment initial data and vague convergence of the initial empirical measures (Demircigil et al., 13 May 2025).

Mean-field interacting branching diffusions with logistic binary branching admit both qualitative and quantitative limits. Under i.i.d. initial atoms, Lipschitz coefficients, and bounded Lipschitz kernels, one has Kt\mathcal K_t3 in Kt\mathcal K_t4, where Kt\mathcal K_t5 is the unique continuous finite-measure solution of

Kt\mathcal K_t6

Its total mass Kt\mathcal K_t7 solves the logistic ODE Kt\mathcal K_t8 (Fontbona et al., 2021). With higher moments, the convergence rate is explicit: Kt\mathcal K_t9 The proof uses a coupling with an auxiliary process of McKean–Vlasov diffusions and optimal transport at branching times (Fontbona et al., 2021).

McKean–Vlasov branching diffusion itself is obtained as a large-population limit. For the μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)0-particle interacting system in which coefficients depend on the empirical law, the law of the empirical measure is tight in μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)1, and any limit point is supported on solutions of the McKean–Vlasov martingale problem. If strong uniqueness holds, the entire sequence converges to the law of the unique McKean–Vlasov branching diffusion (Claisse et al., 2024).

Several interacting branching models admit discrete-to-continuous scaling limits. In the multitype interacting branching framework, if μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)2 is the discrete-state interacting multitype branching process and μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)3 satisfy the scaling assumptions, then

μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)4

where μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)5 is the unique strong solution of the continuous-state interacting branching SDE with quadratic interaction terms (Fittipaldi et al., 2022). In branching processes with competition, the renormalized population size converges to the nonlinear diffusion μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)6, and the whole family μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)7 is represented by local times of a reflected Brownian motion with local-time-dependent drift, yielding a generalized Ray–Knight theorem (Ba et al., 2013).

The finite-system scheme for infinite-rate mutually catalytic branching has a different time scale because the jumps lack finite second moments. For the mean-field site space μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)8, the correct scale for the total masses is

μt=Law(Zt)\mu_t=\mathrm{Law}(Z_t)9

and the rescaled averages tt0 converge to

tt1

with joint convergence of total masses and coordinate processes under the finite-system scheme (Doering et al., 2015).

4. Traveling waves, invasion speeds, and genealogical front classification

For the rank-based Go or Grow limit PDE, traveling waves are obtained from the ansatz tt2, tt3, with cumulative tail tt4. The resulting ODE admits a minimal wave speed

tt5

such that for each tt6 there is a monotone profile tt7 with tt8 and tt9, while no such wave exists for dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,0 (Demircigil et al., 13 May 2025). The transition occurs at the critical value dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,1. When dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,2, the front is pulled and dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,3 coincides with the linear spreading speed of dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,4. When dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,5, the front is pushed and dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,6 (Demircigil et al., 13 May 2025).

Microscopic finite-dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,7 simulations use the position of the dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,8-th largest particle,

dXtk=b(t,Xtk,μt)dt+σ(t,Xtk,μt)dWtk,dX^k_t = b\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dt + \sigma\bigl(t,X^k_{\cdot\wedge t},\mu_t\bigr)\,dW^k_t,9

and the empirical speed estimator

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},00

For each fixed E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},01, E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},02 grows roughly linearly in E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},03. As E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},04 grows, E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},05 concentrates around a deterministic limit, and numerically

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},06

with small finite-E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},07 corrections in the pulled regime E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},08, reflecting the Bramson logarithmic shift (Demircigil et al., 13 May 2025).

The same model supports a genealogical criterion for pulled versus pushed fronts. If a particle sampled at time E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},09 is followed backward along its unique ancestral lineage, and the lineage is observed in the moving frame of the front, then Conjecture 5.5 states that for large E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},10

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},11

In the pushed case E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},12, the adjoint operator admits a unique integrable invariant density

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},13

and numerically the lineage histogram closely matches E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},14. In the pulled case E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},15, the formal stationary solution is not integrable, so the lineage law drifts to E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},16; ancestors come only from the ever-advancing tip. Exactly at E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},17, a hybrid “pushmi-pullyu” behavior is observed: the mode stays near E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},18, but the mean drifts out, in agreement with the Bramson correction exponent E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},19 (Demircigil et al., 13 May 2025).

A related rank-based branching-selection system also converges to a reaction-diffusion equation. In the E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},20-BBM family, the hydrodynamic limit is

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},21

and under additional assumptions the particle system has an asymptotic velocity

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},22

The minimal spreading speed of the PDE is exactly E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},23, and E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},24, which is described there as a partial weak-selection principle (Mercer, 6 May 2026).

5. Long-time behavior, coexistence, extinction, and regulation

In mutually catalytic branching with negative correlations, the long-time behavior is governed by the transience or recurrence of the migration chain. For each fixed E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},25 and E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},26, coexistence of the two types occurs if and only if the underlying migration chain is transient; equivalently, if

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},27

Otherwise, in the recurrent case,

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},28

In particular, on E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},29, coexistence occurs precisely for E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},30 (Doering et al., 2011).

The discrete particle system with mutually catalytic interactions on E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},31 has the same phase boundary. For nearest-neighbor random walk, E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},32 implies coexistence possible, while E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},33 implies coexistence impossible (Fugenfirov et al., 2023). On finite tori, the empirical total masses converge after renormalization to the mutually catalytic diffusion

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},34

in dimensions E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},35, under the transience bound E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},36 (Fugenfirov et al., 2023).

In branching diffusion with Ornstein–Uhlenbeck drift and pairwise attraction or repulsion, the center of mass decouples from the interaction parameter E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},37. Writing

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},38

one obtains

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},39

Hence, almost surely, E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},40 when E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},41, whereas E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},42 when E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},43, with E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},44 Gaussian (Englander et al., 2016). For the relative system E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},45, if E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},46 then the empirical measure converges almost surely to

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},47

while for E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},48 local extinction always occurs: every fixed bounded set is eventually empty (Englander et al., 2016).

Self-regulation and competition replace coexistence thresholds by carrying-capacity effects. In the multi-type spatial branching model for local self-regulation, the diffusion limit has local logistic drift E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},49, and the abstract states that the densities for large times do not depend on the initial distribution but mainly on the carrying capacities (Greven et al., 2015). In the one-dimensional competition model,

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},50

subcriticality and extinction are characterized by

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},51

(Ba et al., 2013).

These results show that interacting branching diffusions exhibit several distinct long-time regimes: coexistence driven by transience, extinction driven by recurrence or outward drift, regulation by carrying capacities, and front propagation governed by minimal-wave-speed selection.

6. Duality, spine methods, control, and structural representations

Duality is a principal tool when interaction destroys the classical branching property. For negatively correlated mutually catalytic branching, the exact mixed second-moment formula is

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},52

where E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},53 is the collision local time of two independent migration chains (Doering et al., 2011). Since E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},54, the factor E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},55 decays on collisions, which is the key mechanism behind extinction in the recurrent case (Doering et al., 2011). The same paper combines this with exit-time estimates for correlated Brownian motion in the first quadrant and uniform integrability bounds independent of the branching rate, allowing passage to the infinite-rate limit (Doering et al., 2011).

Exponential duality also appears in multitype self-regulating systems. In the exchangeable case, Greven–Sturm–Winter–Zähle construct a dual process E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},56 and the duality function

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},57

which yields uniqueness of the exchangeable diffusion martingale problem and supports equilibrium and coexistence analysis (Greven et al., 2015). For mutually catalytic branching particle systems on finite tori, approximate self-duality based on

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},58

is central in identifying the finite-system-scheme limit (Fugenfirov et al., 2023).

Spine constructions provide another structural method. For continuous type-space branching processes with interactions, Medous introduces a Girsanov-type change of measure that singles out a distinguished lineage and proves the many-to-one identity

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},59

with path-integral weight E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},60 determined by the nonlinear operator E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},61 (Medous, 2023). If E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},62, then E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},63, recovering a classical many-to-one formula. The same framework yields a generalized continuous-time Kesten–Stigum theorem with interactions and an alternative spine construction for exact simulation of stochastic size-dependent populations (Medous, 2023).

A complementary structural representation is the generalized Ray–Knight theorem for branching with competition. Under Hypothesis B, the reflected Brownian motion

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},64

has local times whose inverse-local-time slices encode the diffusion E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},65, and for E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},66,

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},67

This identifies the full branching diffusion field with the local-time profile of a single reflected Brownian motion whose drift depends on accumulated local time through E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},68 (Ba et al., 2013).

Optimal control adds a PDE layer to interacting branching diffusions. In the controlled setting, the dynamic programming principle yields an infinite system of coupled Hamilton–Jacobi–Bellman equations on the disjoint union E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},69, with value functions

E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},70

Under coercivity, continuity and growth bounds transfer to the value function, which is the unique continuous viscosity solution in the corresponding growth class; a comparison principle is proved by a doubling-of-variables argument with a penalization in E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},71 (Ocello, 16 Jan 2026). In the permutation-invariant mean-field regime, optimal controls can be chosen symmetric, and the HJB system reduces to a single PDE on measure space, written in the paper as a master-equation involving non-local derivatives E  =  {λ=iVδ(i,xi):VIc finite,  xiRd},E\;=\;\Bigl\{\lambda=\sum_{i\in\mathcal V}\delta_{(i,x_i)}:\mathcal V\subset\mathcal I_c\text{ finite},\;x_i\in\mathbb R^d\Bigr\},72 (Ocello, 16 Jan 2026).

Taken together, these methods show that branching diffusion with interactions is not a single model but a framework. Its state spaces are measure-valued and genealogical, its nonlinearities may be rank-based, catalytic, mean-field, or competitive, and its analysis proceeds through martingale problems, hydrodynamic limits, scaling limits, traveling-wave theory, duality, spinal decompositions, and viscosity solutions.

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