CRANE: Causal Relevance Analysis of Language-Specific Neurons in Multilingual Large Language Models

Abstract: Multilingual LLMs achieve strong performance across languages, yet how language capabilities are organized at the neuron level remains poorly understood. Prior work has identified language-related neurons mainly through activation-based heuristics, which conflate language preference with functional importance. Prior work has identified language-related neurons mainly through activation-based heuristics, which conflate language preference with functional importance. We propose CRANE, a relevance-based analysis framework that redefines language specificity in terms of functional necessity, identifying language-specific neurons through targeted neuron-level interventions. CRANE characterizes neuron specialization by their contribution to language-conditioned predictions rather than activation magnitude. Our implementation will be made publicly available. Neuron-level interventions reveal a consistent asymmetric pattern: masking neurons relevant to a target language selectively degrades performance on that language while preserving performance on other languages to a substantial extent, indicating language-selective but non-exclusive neuron specializations. Experiments on English, Chinese, and Vietnamese across multiple benchmarks, together with a dedicated relevance-based metric and base-to-chat model transfer analysis, show that CRANE isolates language-specific components more precisely than activation-based methods.